William Bateson (1861-1926) and R. C. Punnett (1875-1967)
Bateson and Punnett, in England, performed experiments on the color of sweet peas and also on the comb shape of chickens. They discovered unusual ratios in these traits and attributed them to the action of two different genes. They also made observations that led to the discovery of the linkage and crossing-over of genes (1905). Bateson postulated the production and transmission of discontinuous variations, and the results on the origin of species (1894).